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The Echoes of Mavolia are the power behind the Thanatos and Grimslies of Mavolia. They are Echoes accumulating within the petrified Mana Tree due to the low levels of Mana in Dawn of Mana which end up possessing the Tree Maidens Anise and Ritzia.

The Door to Mavolia was sealed by Granz following his lover the Tree Maiden Anise's possession by the Echoes and was opened following Ritzia's possession by both the Echoes and Tree Maiden Anise in turn.

They are ultimately destroyed along with Tree Maiden Anise when the Mana Tree is resurrected and Ritzia merges with Faye and the Mana Seed to become a Mana Goddess. Anise herself in a fit of rage proceeded to infect all worlds with her hatred.

The Echoes before their destruction were apparently partially freed(according to Anise herself) when Anise and Ritzia released the Thanatos and as their Darkness in the form of Thanatos spread across the land they manifested as the Benevodons to ravage the land before being ultimately sealed by the Mana Goddess whose ascension destroyed the still sealed remnants of the Echoes that didn't become Benevodons.

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Characters
Main characters
Keldric - Ritzia - Faye
Supporting characters
Buju - Village Elder - Lekius - Chobee - Treant - Gaia - Flammie - Watts
Elemental Spirits
Gnome - Jinn - Salamander - Undine - Wisp - Shade - Luna - Dryad
Villains
Stroud - Masked Guru - Anise
Enemies
Enemies - Bosses
Locations
Fa'Diel - Jadd - Topple - Wendell - Ishe - Lorimar - Illusia Isle - Treefolk Village
Dungeons
Tutorial Area - Great Tree (Chapter 1) - Wonderwood - Mortmont - Imperial Flagship - Jadd Castle - Beacon Tower - Great Tree (Chapter 7) - Nethermanse
Terms
Mana Tree - Mana Sword - Tree Maiden - Grimslie - Tree Folk - Guardian Beast - Echoes - Mavolia - Door to Mavolia - Thanatos - Mark II
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